As Easter 1940 approaches, James tries to help a young boy and his sick dog, Audrey faces her past, while Siegfried misses Tristan.

Siegfried hires an experienced bookkeeper, but her efficient approach doesn't suit everyone. James and Helen arrive at a decision, and a ferret presents a challenge for the practice.

When new vet Richard Carmody joins the practice, James feels his place by Siegfried's side is threatened. Conflicts come to the surface at Mrs Pumphrey's community tea party.

Helen rallies the community to help the new farmers while she waits for some personal news. Carmody meets Bernard the tortoise and tries to treat the owner too.

The day that James and Helen have been dreading arrives. Mrs Hall takes a step forward with Gerald. Carmody finds his place at Skeldale, assisted by a surprising mentor.

Helen decides to move back with her father while James is away. Mrs Hall is trying to find the right time to give notice. Siegfried is frustrated by a constant leaking tap in the kitchen.

Faced with a daunting future, James longs to see Helen, but an injured bird stands in his way. Siegfried and Carmody try to support Mrs Hall's community efforts.